RESEARCH TEAM

Meet the researchers behind Joyful by Nature

The Joyful by Nature project brings together researchers from multiple institutions working across animal cognition, behavioural science and evolutionary biology.

Principal Investigators

Colin Allen

Philosopher and Cognitive Scientist at the University of California, Santa Barbara, USA

Colin Allen

Philosopher and Cognitive Scientist at the University of California, Santa Barbara, USA

Allen publishes on the philosophical foundations of cognitive science with particular emphasis on animal minds and artificial intelligence.

Erica Cartmill

Primatologist and Comparative Psychologist at Indiana University Bloomington, USA

Erica Cartmill

Primatologist and Comparative Psychologist at Indiana University Bloomington, USA

Erica Cartmill is Professor of Cognitive Science, Anthropology, and Animal Behavior at Indiana University. Her research explores the evolution of language and social cognition

Vincent Janik

Marine Biologist and Cognitive Scientist at the University of St Andrews, UK

Vincent Janik

Marine Biologist and Cognitive Scientist at the University of St Andrews, UK

Vincent Janik is a Professor of Marine Biology and Director of the Scottish Oceans Institute at the University of St Andrews. His research focuses on cognition and communication in marine mammals and how anthropogenic noise affects these animals. Janik led numerous marine field expeditions and serves regularly on governmental and non-governmental advisory boards in his field. He teaches students at all levels in marine mammalogy, bioacoustics and animal behaviour.

Heidi Lyn

Comparative Psychologist and Marine Scientist at the University of South Alabama, USA

Heidi Lyn

Comparative Psychologist and Marine Scientist at the University of South Alabama, USA

Studies cognition and communication across species including apes, dolphins, dogs, and hermit crabs

Ximena Nelson

Neuroethologist and Behavioral Ecologist in the School of Biological Sciences at the University of Canterbury, Christchurch, New Zealand

Ximena Nelson

Neuroethologist and Behavioral Ecologist in the School of Biological Sciences at the University of Canterbury, Christchurch, New Zealand

Ximena Nelson specialises in animal behaviour, animal cognition, sensory ecology, and neuroethology. Her research examines how animals perceive, process, and act on information, with a particular focus on kea parrots and jumping spiders, two systems that illuminate the evolution of intelligence, decision‑making, and sensory‑driven behaviour. Her research has been featured in major scientific outlets and international media.
Relevant publications:

Grabham AA, Mill MM, Florent SN, Eriksen M, Prokshina A, Hill J, Thompson D, Taylor AH,

Nelson XJ 2026. A disabled kea parrot is the alpha male of his circus. Current Biology. In Press. Nelson XJ, Taylor AH, Cartmill EA, Lynn H, Robinson LM, Janik V, Allen C 2023. Joyful by nature: Approaches to investigate the evolution and function of joy in non-human animals. Biological Reviews 98, 1548–1563. doi:10.1111/brv.12965.

Alex Taylor

Comparative Psychologist at the Catalan Institution for Research and Advanced Studies, Barcelona, Spain

Alex Taylor

Comparative Psychologist at the Catalan Institution for Research and Advanced Studies, Barcelona, Spain

Alex Taylor works across the fields of comparative psychology, behavioural neuroscience, evolutionary biology, animal behaviour, and animal welfare. The key focus of his research is the development of experimental methodologies and theoretical frameworks that increase the strength of the inferences we can make about the cognitive processes of animals.

Relevant publications: ·

Nelson XJ, Taylor AH, Cartmill, EA, Lyn H, Robinson LM, Janik V & Allen C (2023) Joyful by nature: approaches to investigate the evolution and function of joy in non-human animals. Biological Reviews 98(5), 1548-1563. IF 14.4

· Taylor AH, Bastos APM, Brown R, & Allen C (2022) The signature-testing approach to mapping biological and artificial intelligences Trends in Cognitive Sciences 26(9), 738-750. IF 21.8

Other Personnel

Gal Badihi (Cartmill)

Researcher/Biologist

PhD student at University of St. Andrews

Alex Grabham (Nelson)

Researcher/Biologist

PhD student at University of Canterbury

Tom Jenks (Janik)

Researcher/Biologist

PhD student at University of St. Andrews

Daan Laméris (Cartmill)

Researcher/Biologist

Postdoctoral researcher, UCLA and Antwerp ZOO Centre for Research and Conservation

Lydia McClean (Nelson)

Researcher/Biologist

PhD Student, Canterbury University

Patrick Neilands (Taylor)

Researcher/Biologist

PhD Student, University of Auckland

Patrick Neilands (Taylor)

Researcher/Biologist

PhD Student, University of Auckland

Joclyn Villegas (Lyn)

Researcher/Biologist

PhD Student, University of South Alabama

Lindsey Johnson

Dr. Lindsey

Johnson is a postdoctoral researcher in the C3P0 Lab at the University of South Alabama currently focused on studying positive emotion in apes and dolphins.

Collaborators

Stefano Furlati

Scientific Education Director

Oltremare Marine Park, Riccione,Italy

Catherine Hobaiter

Professor

School of Psychology and Neuroscience, St. Andrews University

Julie Oswald

Senior Lecturer

School of Biology, St. Andrews University

Jill Pruetz

Professor

Department of Anthropology, Texas State University

Research Partners